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The park covers an area of 31,160 hectars characterised by mountains, including Pic Boby, which at 2,658 metres is Madagascar's highest accessible peak.
Andringitra is an important central link in the longest unbroken chain of remaining rainforest in Madagascar, stretching some 180 kilometres from [[Ranomafana National Park ]] in the north to [[Peak Ivohibe]] in the south.
The park is divided into three ecosystems: low altitude rainforest, montane highland forest and high altitude vegetation. Climates range between humid and tropical in the eastern lowland rainforests to extreme cold temperatures at higher elevations, which can descend to -8°C.